Eleutherodactyline Frogs (Anura: Leptodactylidae) from the Cordillera Yanachaga in Central Peru
A small collection of anurans from the Cordillera Yanachaga in central Peru revealed the presence of 10 species of Eleutherodactylus. Four of these species are unknown elsewhere and are described herein. Three of the new species are members of the immense Eleutherodactylus unistrigatus group, and one belongs with the smaller, South American Eleutherodactylus conspicillatus group. The 83 described species of Eleutherodactylus known in Peru belong to six species groups that have different patterns of distribution. Most species in the cloud forest in the Cordillera Oriental, Cordillera Central, and associated ranges have small latitudinal and elevational distributions. Una pequeña colección de anuros de la Cordillera Yanachaga en el centro del Perú ha revelado la presencia de 10 especies de Eleutherodactylus. Cuatro de estas especies son únicas de esta región y se describen en este trabajo. Tres de las especies nuevas pertenecen al inmenso grupo Eleutherodactylus unistrigatus y la especie adicional pertenece al grupo Eleutherodactylus conspicillatus. Las 83 especies de Eleutherodactylus conocidas en el Perú están incluídas en seis grupos de especies que tienen diferentes patrones de distribución. La mayoría de las especies que habitan los bosque nublados de la Cordillera Oriental, Cordillera Central y serranías asociadas presentan rangos de distribución latitudinal y altitudinal restringidos.Abstract
Resumen

Phylogenetic tree of some South American species of Eleutherodactylus and Phrynopus showing relationships of specimens collected in the region of Oxapampa, Peru, including new species described herein. Numbers on nodes are bootstrap confidence values. The tree was rooted with Rana pipiens (not shown). The scale bar indicates percent sequence divergence

New species of Eleutherodactylus. (A) E. bipunctatus, KU 291638, female, 35.5 mm SVL. (B) E. aniptopalmatus, KU 291626, female, 22.0 mm SVL. (C) E. rhabdocnemus, KU 291646, female, 24.5 mm SVL. (D) E. stictogaster, KU 291659, female, 20.5 mm SVL. Photographs by S. Blair Hedges
Contributor Notes
(WED) Natural History Museum and Biodiversity Research Center, University of Kansas, Lawrence, Kansas 66045; and (SBH) Department of Biology, 208 Mueller Laboratory, Pennsylvania State University, University Park, Pennsylvania 16802. (WED) duellman@ku.edu; and (SBH) sbh1@psu.edu Send reprint requests to WED.