Revision of Sauvagella Bertin (Clupeidae; Pellonulinae; Ehiravini) with a Description of a New Species from the Freshwaters of Madagascar and Diagnosis of the Ehiravini
The Madagascan genus Sauvagella is revised and a new species, Sauvagella robusta, from the Ambomboa River (Sofia drainage) described. Preliminary analysis suggests Sauvagella is an ehiravin and that the pellonuline tribe Ehiravini is monophyletic. The immediate relationships of Sauvagella appear to lie with the single other ehiravin genus from Madagascan waters, Spratellomorpha. Reports of the occurrence of Gilchristella in Madagascan waters are in error.Abstract

Sauvagella madagascariensis (A) holotype, MNHN 3795, female 49.7 mm SL. (B) AMNH 231409, female 51.8 mm SL

Upper jaw in lateral view of (A) Sauvagella madagascariensis, AMNH 88142, 52.0 mm SL (B) Sauvagella robusta, AMNH 231419, 44.0 mm SL. Lower limb of first gill arch of (C) S. madagascariensis, AMNH 88142, 52.0 mm SL (D) S. robusta, AMNH 231419, 44.0 mm SL. Hyoid arch in lateral view of (E) S. madagascariensis, AMNH 88142, 52.0 mm SL (F) S. robusta, AMNH 231419, 44.0 mm SL. Suspensorium in lateral view, (G) S. madagascariensis, AMNH 88142, 52.0 mm SL. (H) Palatine, ectopterygoid and endopterygoid of S. robusta, AMNH 231419, 44.0 mm SL

Madagascar (Mangarahara River indicated), with presumed ranges for Sauvagella and Spratellomorpha

Sauvagella robusta (A) holotype, AMNH 231420, female 47.1 mm SL, drawn by Patricia Wynne (B) photograph of same fish

Caudal fin skeleton of (A) Hyperlophus translucidus AMNH 55309, 43.0 mm SL (B) Spratellomorpha bianalis AMNH 231358, 34.0 mm SL (C) Ehirava fluviatilis AMNH 229124, 42.0 mm SL (D) Sauvagella madagascariensis AMNH 88142, 52.0 mm SL

Alternative schemes of ehiravin intrarelationship (A) modified after Grande, 1985, (B) this study

Posterior region of anal fin and associated structures in (A) Spratellomorpha bianalis AMNH 231358, 34.0 mm SL (B) Sauvagella madagascariensis, AMNH 88142, 52.0 mm SL, and (C) Clupeichthys bleekeri, AMNH 48878, 50.0 mm SL